Our research endeavors are aimed at understanding the mechanism of growth in crustaceans and, in particular, cellular processes occurring in crustacean larvae. We are also studying the fouling community in a local estuary and developing methods for treating ship's ballast water to counter the introduction of invasive species.

Research Projects:

  • Role of nutrients in cellular growth and cell differentiation in larval epidermal tissues .
  • Role of highly unsaturated fatty acids in limb growth and development in brine shrimp larvae.
  • Cellular enlargement in epidermal tissues in developing shrimp larvae.
  • Changes in the fouling community in nearby estuarine habitats .


-Freeman John A, 1993. The crustacean epidermis during larval development. In: M.N. Horst and J.A. Freeman (eds) The Crustacean Integument: Morphology and Biochemistry. CRC Press, Boca Raton, FL. Chapter 6.

-Freeman, John A and J.D. Costlow, 1980. The molt cycle and its hormonal control in Rhithropanopeus harrisii larvae. Develop. BioI. 74,479-485.

-Freeman, John A and J.D. Costlow, 1983. The cyprid molt cycle and its hormonal control in the barnacle Balanus amphitrite . J. Crustacean BioI. 3, 173-182.

-Freeman, John A, T. L. West, and J.D. Costlow, 1983. Postlarval growth in juvenile Rhithropanopeus harrisii . BioI. Bull. 105,409-415.

-Freeman, John A, G. Kilgus, D. Laurendeau, and HM. Perry, 1987. Postmolt and intermolt molt cycle stages of Caliinectes sapidus . Aquaculture 61, 201-209.

-Freeman, John A and R.B. Chronister. 1989. Cell cycle kinetics of epidermal cells in developing Artemia : in situ analysis. Journal of Crustacean Biology 9, 193-201.

-Freeman, John A 1990. Molt increment, molt cycle duration, and tissue growth in Palaemonetes pugio Holthuis larvae. J. Expo Mar. BioI. Ecol. 143,47-61.

-Freeman, John A 1990b. Regulation of tissue growth in crustacean larvae by feeding regime. BioI. Bull. 178,217-221.

-Freeman, J.A., L.B. Cheshire, and T.M. MacRae. 1992. Epithelial morphogenesis in developing Artemia : The role of cell replication, cell shape change, and the cytoskeleton. Develop. Biology 152, 279-292.

-Freeman, J.A 1995. Epidermal cell cycle and region-specific growth during segment development in Artemia . J. Exp. Zool. 271, 285-295.

-MacRae, T.H and J.A Freeman. 1995. Organization of the cytoskeleton in brine shrimp setal cells is molt-dependent. Canadian 1 Zoology 73, 765-774.

-Freeman, J.A and M. Porterfield. 1996. Nutrient stimulation of epidermal growth in developing Artemia . Aquaculture Nutrition 2,201-206.

-Perry, H, C. Trigg, K. Larsen, J. Freeman, M. Erickson, and R. Henry. 2001. Calcium concentration in seawater and exoskeletal calcification in the blue crab, Callinectes sapidus . Aquaculture 198, 197-208.

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